Thanks for this, Nearly. I also found it a fascinating and expansive article which echoes some of my own views. It has been so easy to anthropomorphise and sentimentalise animal behaviour and even to see animals in an almost 'Wind in the Willows' like human form, that only in the last 20 to 30 years have accurate observations been made on a range of animal species leading, as the article makes clear,to the view that a variety of human tendencies do seem to be replicated in certain other animal species. I would suggest that technological advances have greatly helped in studying such animal behaviour(e.g. photography, sound recording). I particularly liked his reference to insects(whatever it feels like to be a bee, it feels something). Indeed this short article may interest you, which suggests that even fruit flies seem to be capable of 'free will' whatever that is, of course.
https://www.sciencedaily.com/releases/2007/05/070516071806.htm
